The women of Mulberry Lane anxiously wait for news. Peggy doesn’t know if she’ll ever again see Able, the American father of her twins, while a very odd letter from estranged husband, Laurie, threatens to set the cat among the pigeons. Rose can’t stop loving Jimmy, while Tom can’t stop loving her. Will Gordon ever again be the man he once was? Can he be a proper husband to Maureen, and if he can’t, what will become of their marriage?

When Sally, Beth, Margaret and Rachel meet at a job interview for the wonderful new store in Oxford Street, they have no idea they will become lifelong friends. When all four girls are lucky enough to be selected as sales staff their exciting new adventure begins. Join them as they overcome heartbreak and grief, find love and happiness and remain united in their friendship, whatever life throws at them.

London, 1938: Lizzie Larch is a 20-year-old hat maker in London's east end. She is happy and popular, but she carries a secret. Seven years ago she was viciously attacked and recovered in a private sanatorium where she miscarried a child. Lizzie has no memory of the night of the attack, but secrets cannot stay secret for too long. When she starts courting her boss' nephew, a shocking revelation surfaces; threatening to destroy their new-found happiness.

Cambridgeshire - March 1939. As the clouds of war begin to gather in Europe, the Talbot family of rural Blackberry Farm will be torn apart, just as so many families all over the world will be. Life will never be the same again. Whilst in London, the Salmons family will feel the pain of parting and loss. Brought together by war, the two families become intertwined and, as the outlook looks bleak, they must draw on each other’s strength to fight through the hard times.

As a small child, Nellie Peace was always dreaming but sensed her mother’s rejection. Abandoned and sent into service at Beaumont House at an early age, Nellie is lost and alone until she meets the unpredictable and reclusive artist Lucas Harrington and falls in love with him. This unlikely association between master and servant is encouraged by Lucas’ gentle-natured Aunt Alice, as Lucas sees something unusual in Nellie and is compelled to paint her. Broken promises lead to inevitable heartbreak, and Nellie flees Beaumont House in disgrace for London.

At the heart of Dover's Market Square lies the Castle's Café. A bustling, homely place run by the formidable Nellie Castle and her six children. Since the scandalous birth of her beloved son 10 years before, Marianne, Nellie's eldest daughter, has preferred to stay in the kitchen, hidden away from the town gossips. Overcome with shame, Marianne has never revealed the name of Donny's father - not even to her own mother. But with World War II just around the corner, Marianne's past catches up with her. And suddenly the lives of the Castle family become a lot more complicated.

Alice Lake has arrived in London from Liverpool to start her training as a District Nurse, but her journey has been far from easy. Her parents think that she should settle down and get married, but she has already had her heart broken once and isn’t about to make the same mistake again. Alice and her best friend, Edith, are based in the East End, but before they’ve even got their smart new uniforms on, war breaks out, and Hitler’s bombs are raining down on London.

As Samantha Wishaw watches the love of her life marry another woman, she’s ready to give up hope of finding her happy ever after. But when a chance encounter leads Sam to find friendship in Thomas - a lonely, grumpy elderly widower living at derelict Hedgehog Hollow - her life is about to change forever. Glad to have a distraction from family feuds and match-making, Sam vows to fulfill Thomas and his wife Gwendoline's dreams of restoring Hedgehog Hollow to its former glory, and to open a hedgehog rescue centre.

After traveling the world, Dr. Cody Ryan has finally come home to his foster family’s ranch in Honey Grove, Texas. But all his time in Doctors Without Borders couldn’t have prepared him for the sudden blizzard that forces him to take shelter in an old barn - or for the shock of watching Stephanie O’Dell yank open that same barn door minutes later. He’s barely seen the gorgeous veterinarian since he returned, so why is she icier than the wind outside?

Robinswood, Co. Waterford, 1939. The once grand house is home to two very different families. Despite delusions of grandeur, Lord and Lady Kenefick and their adult children live a life of decayed opulence as the money needed to keep such a large house and grounds ever dwindles. Meanwhile, the Murphy family, Dermot, Isabella and their three almost grown-up girls, live and work on the estate and do their best to keep everything running smoothly.

It's 1942, and World War Two is in full swing. Roberta 'Bobby' Hollis has joined the Air Transport Auxiliary in a team of pilots known as the 'glamour girls'. Just as capable as the men, these women ferry aircraft, parts and people between factories, assembly plants and airfields. 

As the German Luftwaffe begin a terrifying bombing campaign that will come to be known as the Blitz, thousands are evacuated to safety. But Lizzie Mackenzie finds herself heading toward London. She knows she must help in the war effort. But she has another reason for leaving the security of her Scottish village: the illegitimate child she gave up for adoption nearly five years before is somewhere in the city. And - as the bombs rain down - she will stop at nothing to find her and make sure her little girl is safe. 

As the Great War continues to take its toll, headstrong 21-year-old Emily Bryce is determined to contribute to the war effort. She is convinced by a cheeky and handsome Australian pilot that she can do more, and it is not long before she falls in love with him and accepts his proposal of marriage. When he is sent back to the front, Emily volunteers as a “land girl,” tending to the neglected grounds of a large Devonshire estate. It’s here that Emily discovers the long-forgotten journals of a medicine woman who devoted her life to her herbal garden.

Liverpool, 1939. Bella Rogers thought her life was just starting; she can’t wait to get married to her childhood sweetheart, Bobby. But when war is declared and Bobby is called up, his letters stop, leaving Bella heartbroken and alone. Then her family is torn apart by tragedy when her angelic 5-year-old sister dies from a terrible illness. They can barely afford a funeral, but the church is packed with the entire community, devastated for their loss. Grieving for her sister and lost love, Bella finds a job at the Bryant and May factory, making matches.

As Britain stands alone against a monstrous enemy, the inhabitants of Pearl Street face hardship and heartbreak with courage and humour. The war touches each of them in a different way: for Annie Poulson, a widow, it means never-ending worry when her twin boys are called up and sent to France; Sheila Reilly's husband, Cal, faces the terror of U-Boat attacks; Eileen Costello is liberated from a bitter, loveless marriage when her husband is sent to Egypt.